Ray Angelo1 and David E. Boufford2
Harvard University Herbaria
22 Divinity Avenue
Cambridge, MA 02138-2020, USA
1rangelo@oeb.harvard.edu
2david_boufford@harvard.edu
ABSTRACT. Dot maps are provided to depict the distribution at the county level of the taxa of Magnoliophyta: Asteraceae
(corresponding to Flora of North America, Vols. 19, 20, 21) growing outside of cultivation in the six New England states
of the northeastern United States. The maps treat 491 taxa (species, subspecies, varieties, and hybrids, but not forms)
based primarily on specimens in major herbaria of Maine, New Hampshire, Vermont, Massachusetts, Rhode Island, and Connecticut,
with most data derived from the holdings of the New England Botanical Club Herbarium (NEBC). Brief synonymy (to
account for names used in standard manuals and floras for the area) habitat, chromosome information, and common
names are also provided.
Key Words: flora, New England, atlas, distribution, Asteraceae.
This article is the ninth in a series (Angelo and Boufford, 1996, 1998, 2000, 2007, 2010, 2011a, 2011b, 2012) that
present the distributions of the vascular flora of New England
in the form of dot distribution maps at the county level (see Key Map). The
atlas is posted on the internet at http://neatlas.org/
where it will be updated as new information becomes available.
This project encompasses all vascular plants (pteridophytes and
spermatophytes) at the rank of species, subspecies, and variety
growing independent of cultivation in the six New England states.
Hybrids are also included, but forms and other ranks below the level
of variety are not. The dots are based on voucher specimens primarily
in New England herbaria (of colleges, universities, botanical gardens,
and public museums) representing reproducing populations
outside of cultivated habitats. This ninth installment includes the
family in Magnoliophyta: Asteraceae corresponding to the family treated
in Flora of North America, Volumes 19, 20, 21 (Flora of North America
Editorial Committee 2006a, 2006b. 2006c). Of the 491 taxa treated,
223 are not native to the region. Future accounts will treat the
distribution of additional non-monocot angiosperms.
The habitat data are distillations from a variety of sources augmented by our
own field observations. An attempt was made to indicate habitat information as
it applies to a particular taxon in New England rather than to the entire range
of the taxon. Such information is omitted where habitat is not indicated on the
specimen label and where we also lack personal knowledge of the plant in New England.
Omissions of habitat information are for a few introduced taxa and for all hybrids.
It is our hope that these articles will stimulate additional field work to supplement the distributions portrayed
in the maps. The New England Botanical Club herbarium has proven to be the most important resource
for this project. We are eager to receive information on voucher specimens in public herbaria documenting
range extensions and filling county gaps in distributions. Similarly, because the atlas of the New England
flora will be continuously updated as new information becomes available, we are eager to receive notification
of published corrections of cytological information and new, documented chromosome counts for taxa in the
New England flora.
Materials and methods are as outlined in Angelo and Boufford (1996) and at
http://neatlas.org/Intro-Pterid&Gym.html
and are not repeated here.
TAXONOMY AND FORMAT
The taxonomy and nomenclature adopted for this work essentially follow
that of the Flora of North America project in progress, except that families,
genera, and species are arranged alphabetically. The families and their
circumscription do not necessarily reflect current views on relationships
or composition. The Angiosperm Phylogeny Website (Stevens 2001 onwards) should
be consulted for a continuously updated treatment of families and their inclusive genera.
Named and unnamed hybrid taxa are placed alphabetically at the end of the genus in which
they occur. Unnamed hybrids combine the names of the progenitors alphabetically by epithet.
Taxa that are not native to New England are indicated by uppercase text. Unpublished names
are not used, even if publication is pending.
Chromosome numbers are taken from Flora of North America, Volumes 19, 20, 21
(Flora of North America Editorial Committee 2006a, 2006b, 2006c) and from Missouri Botanical Garden's
Index to Plant Chromosome Numbers [website
http://www.tropicos.org/NameSearch.aspx?projectid=9; St. Louis, MO]
Synonymy is provided primarily with respect to names accepted in standard
manuals covering New England published from 1950 onward, including Fernald
(1950), Gleason and Cronquist (1991), and Seymour (1982).
Synonyms have not been provided where the distribution for the synonymized
name does not include New England.
The following list (which includes excluded taxa) will aid readers in finding familiar names that have been transferred to other taxa:
The following species are reported from our area but are excluded for the reasons noted:
ACANTHOSPERMUM AUSTRALE (Loefling) Kuntze [no voucher found for wild
occurrence; reported from Massachusetts]
AMBERBOA MOSCHATA Bunge [no specimen located; reported from Maine in Flora of North America Editorial Committee (2003)]
Antennaria rosea Greene subsp. pulvinata (Greene) R.J. Bayer [no specimen located;
reported from Maine]
ARCTANTHEMUM ARCTICUM (Linnaeus) Tzvelev subsp. POLARE (Hultén) Tzvelev [no specimen located; reported from Massachusetts]
ARTEMISIA DRACUNCULUS Linnaeus (A. GLAUCA Pallas ex Willdenow var. DRACUNCULINA (S. Watson) Fernald) [no specimen located; reported from Massachusetts]
BRICKELLIA GRANDIFLORA (Hooker) Nuttall [no specimen located; reported from Rhode Island]
CARTHAMUS LANATUS Linnaeus [no voucher found for wild occurrence; reported from Massachusetts]
CENTAUREA BOVINA Velenovský [no specimen located; reported from Massachusetts]
CENTAUREA PHRYGIA Linnaeus (C. AUSTRIACA Willdenow) [no specimen located; reported from Vermont]
CENTIPEDA MINIMA (Linnaeus) A. Braun & Ascherson [no voucher found for wild occurrence; reported from Massachusetts]
COTULA AUSTRALIS (Sieber ex Sprengel) Hooker f. [no specimen located; reported from Maine]
CHRYSANTHEMUM X MORIFOLIUM (Ramatuelle) Hemsley (pro species) – (C. INDICUM Linnaeus X C. JAPONICUM Thunberg) [no specimen located; reported from
Massachusetts]
CHRYSOGONUM VIRGINIANUM Linnaeus var. BREVISTOLON G.L. Nesom [no specimen located; reported from Rhode Island]
CIRSIUM CANUM (Linnaeus) Allioni [no specimen located; reported from Massachusetts]
CREPIS PANNONICA (Jacquin) K. Koch [no specimen located; reported from Connecticut]
CRUPINA VULGARIS Persoon ex Cassini [no specimen located; reported from Massachusetts]
EUCHITON INVOLUCRATUS (G. Forster) Anderberg (GNAPHALIUM INVOLUCRATUM G. Forster) [no voucher found for wild occurrence; reported from Massachusetts]
EUPATORIUM CAPILLIFOLIUM (Lamarck) Small [no specimen located; reported from Connecticut and Massachusetts]
Eurybia chlorolepis (E.S. Burgess) G.L. Nesom [specimen from Connecticut at CONN identified as this taxon is judged to be misidentified]
EURYBIA SURCULOSA (Michaux) G.L. Nesom (ASTER SURCULOSA Michaux) [no specimen located; reported from Massachusetts and Connecticut]
FLAVERIA TRINERVIA (Sprengel) C. Mohr [no voucher found for wild occurrence; reported from Massachusetts]
GAILLARDIA ARISTATA Pursh [no voucher found for wild occurrence; reported from Connecticut, Massachusetts and New Hampshire]
HELIANTHUS DEBILIS Nuttall subsp. DEBILIS [no voucher found for wild occurrence; reported from Maine]
HELIANTHUS HIRSUTUS Rafinesque [no specimen located; reported from Connecticut]
HELIANTHUS MICROCEPHALUS Torrey & A. Gray [no voucher found for wild occurrence; reported from Connecticut]
HELIANTHUS OCCIDENTALIS Riddell subsp. OCCIDENTALIS Torrey & A. Gray [no specimen located; reported from Massachusetts]
INULA SALICINA Linnaeus [no specimen located; reported from Massachusetts]
Lactuca floridana (Linnaeus) Gaertner [voucher has been re-identified as L. biennis; reported from Massachusetts]
LACTUCA SALIGNA Linnaeus [no specimen located; reported from Maine and Massachusetts]]
LIATRIS LIGULISTYLIS (A. Nelson) K. Schumann [no specimen located; reported from Connecticut]
LIATRIS SCARIOSA (Linnaeus) Willdenow var. NIEUWLANDII (Lunell) E.G. Voss (L. X NIEUWLANDII (Lunell) Gaiser) [no specimen located; reported from Connecticut and Rhode Island]
LOGFIA MINIMA (Smith) Dumortier (FILAGO MINIMA (Smith) Persoon) [no voucher found for wild occurrence; reported from Massachusetts]
PEREZIA MULTIFLORA (Bonpland) Lessing subsp. SONCHIFOLIA (Baker) Vuilleumier (P. ALETES J.F. Macbride) [no voucher found for wild occurrence; reported from Massachusetts]
PETASITES JAPONICUS (Siebold & Zuccarini) Maximowicz [no voucher found for wild occurrence; reported from Maine]
Rudbeckia laciniata Linnaeus var. bipinnata Perdue [no specimen located; reported from Connecticut, Massachusetts and New Hampshire]
SANVITALIA PROCUMBENS Lamarck [no voucher found for wild occurrence; reported from Vermont]
SCHKUHRIA PINNATA (Lamarck) Kuntze ex Thellung [no voucher found for wild occurrence; reported from Maine and Massachusetts]
Senecio pseudoarnica Lessing [no specimen located; reported from Maine]
SENECIO SPARTIOIDES Torrey & A. Gray (S. MULTICAPITATUS Greenman) [no specimen located; reported from Massachusetts]
SERRATULA TINCTORIA Linnaeus [no specimen located; reported from Connecticut]
SIGESBECKIA ORIENTALIS Linnaeus [no specimen located; reported from Middlesex Co., Massachusetts, almost certainly based on the voucher for S. JORULLENSIS Kunth from the same county]
Solidago erecta Banks ex Pursh [no specimen located; specimen from Plymouth Co., Massachusetts at CONN identified as this taxon is judged to be misidentified; reported from Connecticut, Massachusetts and Rhode Island]
SOLIDAGO LEPIDA de Candolle var. SALEBROSA (Piper) Semple (S. CANADENSIS Linnaeus var. SALEBROSA (Piper) M.E. Jones) [no specimen located; reported from Maine]
Solidago X beaudryi B. Boivin (S. rugosa Miller var. rugosa X S. uliginosa Nuttall) [no specimen located; reported from Maine]
Symphyotrichum X gravesii (E.S. Burgess) G.L. Nesom (pro species) -- (S. dumosum (Linnaeus) G.L. Nesom X S. laeve (Linnaeus) Á. Löve & D. Löve var. laeve; Aster gravesii
E.S. Burgess) [no specimen located; reported from Connecticut]
SYMPHYOTRICHUM LANCEOLATUM (Willdenow) G.L. Nesom var. HIRSUTICAULE (Semple & Chmielewski) G.L. Nesom (ASTER LANCEOLATUS Willdenow var. HIRSUTICAULIS Semple & Chmielewski) [no specimen located; reported from
Massachusetts, apparently from incorrect reading of a checklist]
Symphyotrichum novi-belgii (Linnaeus) G.L. Nesom var. crenifolium (Fernald) Labrecque & Brouillet (Aster crenifolius (Fernald) Cronquist) [no specimen located; reported from Maine, New Hampshire and Vermont]
Symphyotrichum phlogifolium (Muhlenberg ex Willdenow) G.L. Nesom (Aster patens Aiton var. phlogifolius (Muhlenberg ex Willdenow) Nees) [no specimen located; reported from Connecticut, Massachusetts, and Rhode Island]
Taraxacum ceratophorum (Ledebour) de Candolle [no specimen located; reported from Massachusetts and New Hampshire]
VERNONIA GLAUCA (Linnaeus) Willdenow [no specimen located; reported from Massachusetts]
ACKNOWLEDGMENTS. We thank the curators and directors of the herbaria of the New England Botanical Club, the Harvard University Herbaria, the University of Massachusetts, and the University of Vermont for allowing access to their collections. For the University of Maine and the University of Connecticut herbaria we used their exceptional online databases of specimens (http://herbaria.umaine.edu/ and http://bgbaseserver.eeb.uconn.edu/databasesimple.html). We are grateful also to Karen Searcy and Roberta Lombardi for facilitating access to the herbarium and to the notebooks of Harry E. Ahles at the University of Massachusetts (Amherst) and to Dr. Searcy for kindly answering requests for information after our visit. James Hinds generously checked information on voucher specimens at the University of Maine (Orono). The following persons also checked certain records for us at their respective institutions: Janet Sullivan, Emily Wood and Elizabeth Allen. In particular we thank the following individuals for repeated checking of specimens at their institution: Robert Capers of the University of Connecticut, Craig D. Layne of Dartmouth College, and Alina Freire-Fierro of The Academy of Natural Sciences of Philadelphia. James Morefield kindly provided information that aided in locating voucher specimens. John Kartesz and Misako Nishino generously provided the latest draft of the Floristic Synthesis of North America, which was consulted for reports of occurrence and sources of such reports. We thank Kanchi Gandhi for nomenclatural assistance.
Top of page
http://neatlas.org/Neatlas8/Intro-Asteraceae.html
-- Revised: Jan. 3, 2013
Created by: Ray Angelo
rangelo@oeb.harvard.edu